The most common variant of the series M1919 was the M1919A4. Production blueprints of the new variant were complete in late 1936, and slow-scale production soon followed. The driving force behind the development of this variant was the suffering reliability of the 18-inch barrel of previous versions, which did not produce enough recoil to cycle the action reliably. The bull barrel was made much thicker and was lengthened to 24 inches like the M1917. Various other small adjustments to the design were made, such as moving the front sight from the barrel jacket to the receiver, which made it easier to mount the gun on vehicles. The design of the barrel jacket was changed to include circular holes instead of long slits of earlier models, and a recoil booster in the muzzle end improved reliability. The recoil buffer assembly was also a new addition to the design between A3 and A4 development, designed to reduce the impact of the bolt hitting the backplate.
The M1919A4 was used in both fixed and flexible mounts, by infantry and on vehicles. When the gun was ready to fire, a round would be in the chamber and the bolt and barrel group would be locked together, with the locking block at the rear of the bolt. When the rear of the trigger was pivoted upwards by the operator, the front of the trigger tipped downward, pulling the sear out of engagement with the spring-loaded firing pin, allowing it to move forward and strike the primer of the cartridge. Two variants were developed specifically for vehicular use, the M1919A5, with an extended charging handle, and the M1919A4E1, a sub-variant of the M1919A4 refitted with an extended charging handle.
The M1919A4 weighed about 14 kg, and was ordinarily mounted on a lightweight, low-slung tripod for infantry use. Fixed vehicle mounts were also employed. It saw wide use in World War II mounted on jeeps, half-tracks, armored cars, tanks, amphibious vehicles, and landing craft. The M1919A4 played a key role in the firepower of the World War II U.S. Army. Each infantry company normally had a weapons platoon in addition to its other organic units. The presence of M1919A4 weapons in the weapons platoon gave company commanders additional automatic fire support at the company level, whether in the assault or on defense.
The M1919A5 was an adaptation of the M1919A4 with a forward mounting point to allow it to be mounted in tanks and armored cars. This, along with the M37 and the Browning M2 machine gun, was the most common secondary armament during World War II for the Allies. The coaxial M37 variant had the ability to feed from either the left or the right of the weapon, and featured an extended charging handle similar to those on the M1919A4E1 and A5. A trial variant fitted with special sighting equipment was designated M37F.



















